born June 6, 1918, Lansing, Iowa, U.S.
U.S. biochemist.
He received his medical degree from Washington University. With Edmond H. Fischer (b. 1920), he won a 1992 Nobel Prize for the discovery of reversible protein phosphorylation, a biochemical process that regulates the activities of proteins in cells and governs countless processes necessary for life. Errors in protein phosphorylation have been implicated in such diseases as diabetes, cancer, and Alzheimer disease.