IRIDACEAE

Crocus. the iris family of flowering plants, belonging to the order Liliales. The family is known for ornamental genera such as Iris, Gladiolus, and Crocus (see photograph). The Iridaceae contains some 80 genera and an estimated 1,700 species of mostly perennial herbs; there are a few shrubs and evergreen herbs as well. The family is nearly worldwide in distribution, but it is most abundant and diversified in Africa. Most species are native to temperate, subtropical, and tropical regions. Species of Iris, Gladiolus, and Crocus grow best in rich soil. A few irises, however, grow in swampy locations, and a few withstand the rigours of subarctic substrates. In addition to Iris, Gladiolus, and Crocus, the Iridaceae includes ornamental genera such as the blackberry lily, or leopard flower (Belamcanda), a native of China and Japan, and the blue-eyed grasses (Sisyrinchium), which are widely dispersed in the Western Hemisphere. The capsule wall of the blackberry lily recurves at maturity to display shiny, black seeds resembling the fruit of the blackberry. The delicate blue-eyed grasses are attractive plants for naturalistic gardens. Plants of the genus Freesia are widely grown commercially for cut flowers. Members of Iris also yield orrisroot (a substance used in the manufacture of perfumes, soaps, powders, and dentrifices). The feathery stigmas of Crocus sativa yield saffron, which is used as flavouring and food colouring and as a medicinal ingredient. The underground stems of members of the iris family may be one of at least three structural types: rhizomes, bulbs, and corms. In Iris the stem is horizontal, robust, and ringed with leaf-scars. It is a rhizome that often grows partially exposed but is firmly rooted in the soil. Species of Iris native to southwestern Europe produce bulbs. This type of stem is short and conical and from it many leaf bases arise, one inside the other. These bases are seamless and constitute the bulk of the bulb. Bulblets arise from the stem, between the leaf bases, to propagate the plant. The corm of Gladiolus is an underground stem in the shape of a doughnut without a hole. On the concave upper surface a small cluster of leaves is located. Among the leaf bases, cormlets arise to reproduce the plant. Rhizomes, bulbs, and corms are efficient in the vegetative reproduction of the Iridaceae. Reproduction by seeds is also very important in the iris family and is essential in the development of new hybrids. In Crocus only one flower may develop from each corm, but in many other genera, as in Iris and Gladiolus, an inflorescence (flower cluster), sometimes branched, arises from the underground stem. Flowers of the showy garden irises possess three sepals (falls), three petals (standards), and three broad, pollen-receptive stigma branches, under which the pollen-producing anthers are hidden. These flower parts are located above the ovary (inferior ovary), which consists of three carpels unified into a single pistil. Ovules within the ovary portion become seeds, and the ovary matures into fruit. Members of the Iridaceae produce dry capsules, a fruit that splits open to release the seeds. Most plants in the family have long, narrow leaves, generally with parallel venation. Bulbous irises may be stored under carefully controlled temperatures, and, when potted, they are brought into flower in cooler months of the year. Temperature also controls the opening of the crocus flower. When near the critical temperature, the perianth (sepals and petals) opens with an increase of less than 1 F. Insects are the pollinators in the iris family, attracted by the showy flower parts. In some gladioli a coordination exists between certain moths and the shape of the flower. While hovering, the moth can reach to the base of the floral tube with its long tongue. Pollen already adhering to the body of the moth is left on the stigma, and fresh pollen sticks to the moth to be left in another flower.