born Oct. 12, 1865, Manchester, Eng.
died June 17, 1940, Bourne, Buckinghamshire
British biochemist.
His more than 20 years of study of sugar fermentation advanced knowledge of metabolic processes in all living forms. In 1929 he shared the Nobel Prize for Chemistry with Hans von Euler-Chelpin. He also produced pioneering studies of bacterial enzymes and metabolism. He was knighted in 1936.