born Aug. 22, 1889, Salem, Va., U.S.
died Oct. 5, 1969, Fort Stockton, Texas
U.S. zoologist and cell biologist.
He received his Ph.D. from Yale in 1913. He was the first to show that the unusually large chromosomes in the salivary glands of Drosophila were particularly well suited for studies of genes and chromosomes. In the early 1930s he published a drawing of a section of a Drosophila chromosome showing more than 150 bands, which for the first time allowed the precise positions of genes to be determined.