MOSS

(class Bryopsida, or Musci), any of at least 10,000 species of small, spore-bearing land plants (division Bryophyta) distributed throughout the world except in salt water. Valvate mosses constitute the subclass Andreaeidae, and peat mosses compose the subclass Sphagnidae. The large subclass Bryidae constitutes most species of mosses, but the subclass Polytrichidae also has some important members. Other, smaller subclasses are represented by only a few species. (See bryophyte.) Mosses are commonly found in moist, shady locations and are best known for those species that carpet woodland and forest floors. Mosses may range in size from microscopic forms to plants more than 1 m (40 inches) long. They differ primarily in the structure and specialization of their sporangia (spore cases). The stemlike and leaflike structures of moss plants constitute the gametophytic (sexual) generation. The sporophytic (asexual) generation develops from the gametophyte and usually consists of a raised stalk, or seta, which terminates in the sporangium. The sporangium remains dependent on the gametophyte, to varying degrees, for water and nutrients. Mosses reproduce by branching and fragmentation, by regeneration from tiny pieces of leaves or stems, and by the production of spores. The spore, under favourable conditions, germinates and grows into a branching, green thread (protonema). Ultimately, the gametophyte grows from an apical cell of the protonema. Mosses break down exposed substrata, releasing nutrients for the use of more complex plants that succeed them. They aid in soil-erosion control by providing surface cover and absorbing water, and they are important in the nutrient and water economy of some vegetation types. Economically important species are those in the genus Sphagnum that form peat. Many small plants bearing the name moss are in fact not mosses. The moss found on the north side of trees is the green alga Pleurococcus; Irish moss is a red alga. Beard lichen (beard moss), Iceland moss, oak moss, and reindeer moss are lichens. Spanish moss is a common name for both a lichen and an air plant of the pineapple family (Bromeliaceae). Club moss is an evergreen herb of the family Lycopodiaceae. Mosses existed as early as the Permian period (286 to 245 million years ago), and more than 100 species were present during the Tertiary period (66.4 to 1.6 million years ago). Muscites, Protosphagnum, Palaeohypnum, and other fossil mosses are similar in structure to modern genera.